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Ulrich Reichard

Ulrich Reichard, Associate Professor

My empirical work focuses on the primate community of Khao Yai National Park, Thailand, where my students and I study white-handed gibbons (Hylobates lar), pileated gibbons (Hylobates pileatus), and northern pig-tailed macaques (Macaca leonina). We continuously monitor individual life histories and collect demographic data. Specific current projects on white-handed gibbons include: behavioral responses to simulated predators, population-level relatedness, sleeping patterns, individuality of song repertoires, and spatial intelligence.




I am interested in relationships between ecology, behavior, and cognition in primates from an evolutionary perspective.

My research is comparative in nature using nonhuman primates as a model to provide data and answers to questions about the evolutionary pressures that have shaped us, humans.

  • ANTH 240A Human Biology: An Introduction to Biological Anthropology
  • UHON 351 Evolution of the Human Mind
  • UHON 351 A Clockwork Nightmare- Life in Nazi Germany
  • ANTH 440C Context of Human Evolution
  • ANTH 455B Laboratory Methods in Biological Anthropology: Modern Field and Laboratory Methods in Wild Animal and Human Studies
  • ANTH 455C Primate Behavior and Ecology
  • ANTH 536 Biology of Desire
  • ANTH 536 Evolution of Primate Intelligence
  • ANTH 536 Primate Cognition, Communication, & Culture
  • ANTH 536 Primate Nutritional Ecology
  • ANTH 518 Socioecology of Apes

In collaboration with Dr. Claudia Barelli (, I direct the field research on white-handed gibbons of Khao Yai, Thailand. The population has been subject to longitudinal research for nearly three decades and the project is ongoing. To facilitate research we maintain a small field station where students and volunteers are welcome to acquire field work experience or collect data for Master’s and Ph.D. degrees. Over the years, more than a dozen groups of gibbons have habituated to the presence of human observers, which allows us to collect data across many social units and individuals. We accept talented, highly motivated students and volunteers to working with us through SIU Carbondale, the German Primate Center (DPZ) or in collaboration with other institutions. For more information contact


Jackson C.P.; Reichard, U.H. (2021). Pygmy marmoset exudate feeding stimulates exudate production. Folia Primatologica 92: 175-182 DOI:10.1159/000516492.

Trébouet F.; Malaivijitnond S.; Reichard, U.H. (2021). Reproductive seasonality in wild northern pig-tailed macaques (Macaca leonina). Primates 62: 491-505.

Andrieu, J.; Penny S.G.; Malaivijitnond, S.; Bouchet, H.; Reichard, U.H.; Zuberbühler, K. (2020). White-handed gibbons (Hylobates lar) discriminate context-specific song compositions. PeerJ. 8:e9477 DOI 10.7717/peerj.9477

Florkiewicz, B.; Skollar, G., Reichard, U.H. (2018). Facial expressions and pair bonds in hylobatids. American Journal of Physical Anthropology 167: 108-123, DOI: 10.1002/ajpa.23608.

Matsudaira, K.; Ishida, T.; Malaivijitnond, S; Reichard, U.H. (2018). Short dispersal distance of males in a wild white-handed gibbon (Hylobates lar) population. American Journal of Physical Anthropology, DOI: 10.1002/ajpa.23603, 167: 61-71.

Terleph T.A.; Malaivijitnond S.; Reichard, U.H. (2018). An analysis of white-handed gibbon male song phrases reveals speech-like syllables. American Journal of Physical Anthropology, DOI: 10.1002/ajpa.23451, 166: 649-660.

Terleph T.A.; Malaivijitnond S.; Reichard, U.H. (2018) Male White-handed gibbons flexibly time duet contributions. Behavioral Ecology and Sociobiology, DOI 10.1007/s00265-017-2432-z, 72:16.

Trébouet F.; Reichard, U.H.; Nantasak P., Malaivijitnond S. (2017). Extractive foraging of toxic caterpillars in wild northern pig-tailed macaques (Macaca leonina). Primates DOI 10.1007/s10329-017-0638-3.

Terleph, T.A., Malaivijitnond S., & Reichard, U.H. (2016). Age related decline in lar gibbon great call performance suggests that call features correlate with physical condition. BMC Evolutionary Biology 16:4 DOI 10.1186/s12862-015-0578-8.

Clarke, E.; Reichard, U.H. & Zuberbühler, K. (2015). Context-specific close-range ”hoo“ calls in wild gibbons (Hylobates lar). BMC Evolutionary Biology 15:56 DOI 10.1186/s12862-015-0332-2.

Terleph, T.A.; Malaivijitnond, S.; Reichard, U.H. (2015). Lar Gibbon (Hylobates Lar) great call reveals individual caller identity. American Journal of Primatology 9999:1-11.

Asensio, N.; Brockelman, W.Y.; Malaivijitnond, S.; and Reichard, U.H. (2014). White-handed Gibbon (Hylobates lar) core area use over a short-time scale. Biotropica 46(4): 461-469.

Reichard, U.H. (2014). The graceful Asian ape. In: Primate Ethnographies, edited by Karen B. Strier. Routledge, New York pp. 118-130.

Barelli, C.; Matsudaira, K.; Wolf, T.; Roos, C.; Heistermann, M.; Hodges, K.; Ishida, T.; Malaivijitnond, S.; and Reichard, U.H. (2013). Extra-pair paternity confirmed in wild white-handed gibbons. American Journal of Primatology 9999: 1-11.

Laurance, W.F. et al. (2012). Averting biodiversity collapse in tropical forest protected areas. Nature 489: 290-294. 

Matsudaira, K.; Reichard, U.H.; Malaivijitnond, S. & Ishida, T. (2012). Molecular evidence for the introgression between Hylobates lar and H. pileatus in the wild. Primates DOI 10.1007/s10329-012-0323-5 

Clarke, E.; Reichard, U.H. & Zuberbühler, K. (2012). The anti-predator behaviour of wild white-handed gibbons (Hylobates lar). Behavioral Ecology Sociobiology 66: 85-96. 

Reichard, U.H.; Ganpanakngan M; and Barelli, C. (2012). White-handed gibbons of Khao Yai: social flexibility, reproductive strategies, and a slow life history. In: Long-term Field Studies of Primates, edited by P.M. Kappeler & D. Watts. Springer, Berlin, pp. 237-258. 

Barelli, C.; Reichard, U.H. & Mundry, R. (2011). Is grooming used as a commodity in wild white-handed gibbons, Hylobates lar? Animal Behaviour 82: 801-809. 

Asensio, N.; Brockelman W.Y.; Malaivijitnond S. & Reichard, U.H. (2011). Gibbon travel paths are goal-oriented. Animal Cognition 14: 395-405. 

Beaman, B.; Hesemeyer, W.; Dominy, N.J.; Savini, T. & Reichard, U.H. (2009). Sterile pyuria in a population of wild white-handed gibbons (Hylobates lar). American Journal of Primatology 71: 880-883.

Reichard, U.H. (2009). Social organization and mating system of Khao Yai white-handed gibbons, 1992 – 2006. In:Wild Gibbon Populations: New Understandings of Small Ape Socioecology, Population Biology and Conservation, edited by S. M. Lappan & D. Whittaker. Springer, Berlin, pp. 347-384. 

Savini, T.; Boesch, C. & Reichard, U.H. (2009). Varying ecological quality influences the probability of polyandry in white-handed gibbons (Hylobates lar). Biotropica 41: 503-513.

Barelli C, Boesch C, Heistermann M and Reichard UH (2008). Female white-handed gibbons (Hylobates lar) lead group movements and have priority of access to food resources. Behaviour 145: 965-981

Barelli C, Heistermann M, Boesch C and Reichard UH (2008). Mating patterns and sexual swellings in pair-living and multimale groups of wild white-handed gibbons (Hylobates lar). Animal Behaviour 75: 991-1001

Reichard UH and Barelli C (2008). Life history and reproductive strategies of Khao Yai white-handed gibbon females (Hylobates lar). International Journal of Primatology 29: 823-844 

Savini T, Boesch C and Reichard UH (2008). Home-range characteristics and the influence of seasonality on female reproduction in white-handed gibbons (Hylobates lar) at Khao Yai National Park, Thailand. American Journal of Physical Anthropology 135: 1-12

Barelli C, Heistermann M, Boesch C and Reichard UH (2007). Sexual swellings in wild white-handed gibbon females (Hylobates lar) indicate the probability of ovulation. Hormones and Behavior 51: 221-230 

Clarke EA, Reichard UH and Zuberbühler K (2006). The syntax and meaning of wild gibbon songs. PLoS ONE 1: e73. doi:10.1371/journal.pone.0000073

Chambers K, Reichard UH, Möller A, Nowak K and Vigilant L (2004). Cross-specific amplification of human microsatellite markers using non-invasive samples from white-handed gibbons (Hylobates lar). American Journal of Primatology 64: 19-27 

Reichard UH (2003a). Monogamy: Past and Present. In: Monogamy Mating Strategies and Partnerships in Birds, Humans and other Mammals (UH Reichard and C Boesch, eds). Cambridge, Cambridge University Press, pp 3-25 

Reichard UH (2003b). Social Monogamy in Gibbons: the Male Perspective. In: Monogamy Mating Strategies and Partnerships in Birds, Humans and other Mammals (UH Reichard and C Boesch, eds). Cambridge, Cambridge University Press, pp. 190-213 

Reichard UH and Boesch C, eds (2003). Monogamy Mating Strategies and Partnerships in Birds, Humans and other Mammals. Cambridge, Cambridge University Press, pp. 276.

Sommer, V. & Reichard, U. (2000). Rethinking Monogamy: The Gibbon Case. In: Primate Males (P. Kappeler, ed.). Cambridge, Cambridge University Press, pp. 159-168.

Reichard, U. (1998). Sleeping Sites, Sleeping Places, and Presleep Behavior of Gibbons (Hylobates lar). American Journal of Primatology, 46: 35-62. 

Brockelman, W.Y.; Reichard, U.; Treesucon, U. & Raemaekers, J.J. (1998). Dispersal, Pair Formation and Social Structure in Gibbons (Hylobates lar). Behavioural Ecology and Sociobiology, 42: 329-339. 

Reichard, U. & Sommer, V. (1997). Group Encounters in Wild Gibbons (Hylobates lar): Agonism, Affiliation, and the Concept of Infanticide. Behaviour 134: 1135-1174. 

Reichard, U. (1995c). Extra-pair Copulations in a Monogamous Gibbon (Hylobates lar). Ethology, 100: 99-112.

Reichard, U. & Sommer, V. (1994). Grooming Site Preferences in Wild White-handed Gibbons (Hylobates lar). Primates. 35: 369-374.

Ulrich Reichard

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